вики ссылается на George Schaller-a: "evidence suggests that the majority die violently from humans or other lions. Lions often inflict serious injuries on each other, either members of different tribes encountering each other in territorial disputes, or members of the same tribe fighting at a kill. Crippled lions and lion cubs may fall victim to hyenas, leopards, or be trampled by buffalo or elephants, and careless lions may be maimed when hunting prey."